The greater importance and vulnerability of the downward flow
of enriched sap, as contrasted to the upward water flow, are apparent.
The phloem conduits are much thinner than the xylem conduits,
and more exposed. Their burden is richer, containing all the tree's
elaborated food, not just raw materials, as in the water column.
External injury to the tree's cambium layer is thus much more
serious than internal injury, to sapwood or heartwood. Trees even
lightly "girdled"—cut or constricted all the way around—
will die, not from the tops down, but from the bottoms up. Deprived
of nourishment from above, the roots wither and cease sending
up water to start the alimentary process.
Exceptional in this respect are palm trees, whose trunks can
suffer circumference damage up to their breaking point without
the trees' health diminishing. This is because the palm family's
phloem conduits are arranged in scattered bundles throughout the
stem instead of in a circle around it.
None of the moisture carried downward in the sapstream to the
roots is returned into the soil. But in nature's economy, trees
do reciprocate earth's gift of water by holding soil, and thus
moisture, in place with their root meshes, and by lessening ground
evaporation with their shade. This is why trees are planted around
reservoirs, to check erosion and parching. Evergreens are most
used for this purpose because they will grow fastest and densest
with the least water requirement for themselves, and their roots
run nearer the surface, where erosion begins.